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Redescription of the male Protoiurus kadleci

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When Protoiurus kadleci (Kovarik, Fet, Soleglad & Yagmur, 2010) (Iuridae) was originally described, the authors thought that they were describing a sexual mature adult. Yagmur and co-workers have now examined more males, and concluded that the original male described was subadult. In the new article, sexually mature males and their hemispermatophores are described.

An updated identification key for the genus is also presented.

Abstract:
Additional material of a rare scorpion species Protoiurus kadleci (Kovařík et al., 2010) from Turkey (Antalya Province) was examined, in particular, sexually mature males. New information is presented on the morphometric differences between adult and subadult males of this species, and on the differences between P. kadleci and other species of Protoiurus. An updated diagnosis of P. kadleci is presented, as well as updated key to six species of Protoiurus, and a map showing all known localities.

Reference:
Yagmur EA, Kovarik F, Fet V, Soleglad ME, Yesilyurt F. Etudes on iurids, IX. Further Analysis of a Rare Species Protoiurus kadleci (Scorpiones: Iuridae) from Turkey, Based on Adult Males. Euscorpius. 2015 (201):1-18. [Open Access]

Family Iuridae

A new species of Diplocentrus from Honduras

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Kevin Sagastume-Espinoza and co-workers have recently published a new species of Diplocentrus Peters, 1861 (Scorpionidae) from Cayos Cochinos in north Honduras, and from a nearby island locality of Utila.

Diplocentrus insularis Sagastume-Espinoza, Longhorn & Santibanez-Lopez, 2015

 An identification key for the genus in Honduras is included.

Abstract:
Three species of genus Diplocentrus are found in north-northwestern Honduras. These species represent the southern east limits of Diplocentrus’ distribution. In recent years, a broad survey of arachnids in Honduras has yielded a collection of several specimens of an undescribed species from two islands in northern Honduras. This new species represents the second species of the genus inhabiting an island. The present contribution describes this new species, and compares it against its most similar relatives. A dichotomous key for the identification of the species of Diplocentrus in Honduras is also included.

Reference:
Sagastume-Espinoza KO, Longhorn SJ, Santibanez-Lopez CE. A new scorpion species of genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) endemic to Islas de la Bahia, Honduras. C R Biol. 2015 May 27. [Subscription required for full text]

Family Scorpionidae

New species in the enigmatic genus Palaeocheloctonus from northern Madagascar

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As many of you already know, Madagascar is a hotspot for scorpion diversity and endemism. Wilson Lourenco and Lucienne Wilme have now described a new species in the enigmatic genus Palaeocheloctonus Lourenco, 1996 (Hormuridae) in northern Madagascar, far from the locations known for the other species in the genus.

Palaeocheloctonus septentrionalis Lourenço & Wilme, 2015

The new species is an indication for a micro-endemic and vicariant population within Madagascar. 

Abstract:
A new scorpion species, Palaeocheloctonus septentrionalis sp.n., is described from the North of Madagascar. The new species clearly suggests a new case of micro-endemic and vicariant population within the island. The number of Palaeocheloctonus species in Madagascar is now increased to two and the known geographical distribution of the genus is extended within that great island, but remains strongly disrupted.

Reference:
Lourenco WR, Wilme L. Micro-endemic populations of Palaeocheloctonus Lourenço, 1996 (Scorpiones: Hormuridae) in Madagascar: A new case of vicariance among Malagasy scorpions. Arthropoda Selecta. 2015;24(2):189-95. [Open Access]

Thanks to professor Lourenco for sending me this paper!

Family Hormuridae

Scorpions of the Galapagos Islands

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The Galapagos Islands are famous for their animals and their impact on our knowledge on animal evolution. But there are also scorpions present on most of the islands in the Galapagos archipelago. Baert & Mahnert have recently published a paper on the scorpion (and other non-spider) fauna of the islands.

Two species are present: Centruroides exsul (Meise, 1934) (Buthidae) and Hadruroides galapagoensis Maury 1975 (Carabotonidae).

Abstract:
The geographic and ecological distribution of the arachnid species belonging to the Amblypygi (Charinus insularis Banks, 1902), the Opiliones (Galanomma microphthalma Juberthie, 1970), the Schizomida (Schizomus portoricensis (Chamberlin, 1922)), the Scorpiones (Centruroides exsul (Meise, 1934) and Hadruroides galapagoensis Maury, 1975), the Solifugae (Neocleobis solitarius (Banks, 1902)) and 25 species of Pseudoscorpiones from the Galàpagos are described. Only the schizomid Schizomus portoricensis and the pseudoscorpions Paratemnoides nidificator (Balzan, 1888), Lechytia chthoniiformis (Balzan, 1887), Aphelolpium cayanum Muchmore, 1979, and Aphelolpium longidigitatum (Ellingsen, 1910) occur also on the American mainland. The pseudoscorpion Withius piger (Simon,1878) is a cosmopolite species.

Reference:
Baert L, Mahnert V. The distribution of the non‐araneae and non‐acari arachnids of Galápagos. Belgian Journal of Entomology. 2015;28:1-76. [Open Access]

Thanks to Rolando Teruel for sending me this article!

Medical important scorpions in Western Brazilian Amazon

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It is well known that scorpions are a major public health problem in many regions of Brazil, but inside Brazil this problem have partly been neglected. But with with an increasing number of sting registrations every year in the country and 78,091 cases reported in 2013, it is important with quality research identifying dangerous species and risk factors.

Amanda M. Queiroz and co-workers have now published a case-controlled study analysis the severity of scorpion stings in Western Brazilian Amazon. Dangerous species are Tityus obscurus (Gervais, 1843), T. metuendus Pocock, 1897 and T. silvestris Pocock, 1897 (family Buthidae). Six deaths were registered and the main conclusion of the study is that scorpion stings showed a extensive distribution in the Western Brazilian Amazon and represent a potential occupational health problem for rural populations in this region.

Abstract:


Reference:
Queiroz AM, Sampaio VS, Mendonca I, Fe NF, Sachett J, Ferreira LC, et al. Severity of Scorpion Stings in the Western Brazilian Amazon: A Case-Control Study. PLoS One. 2015;10(6):e0128819. [Open Access]

The history of three important Tityus species in Brazil

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Professor Wilson Lourenco has recently published an interesting article where he present the history of three important Brazilian Tityus species (Buthidae): Tityus bahiensis (Perty, 1833), Tityus serrulatus Lutz & Mello, 1922 and Tityus costatus (Karsch, 1879).

Abstract:
In the present study, comments are proposed on historical aspects of the most conspicuous scorpion species of the genus Tityus found in Brazil. Both Tityus bahiensis (Perty) and Tityus serrulatus Lutz & Mello are better known for their infamous reputation of noxious species. However, the original discovery and description of both species are associated with interesting historical episodes. A short comment is also provided on Tityus costatus (Karsch), the species possibly involved in the first record of a scorpion incident in Brazil.

Reference:
Lourenco WR. What do we know about some of the most conspicuous scorpion species of the genus Tityus? A historical approach. J Venom Anim Toxins Incl Trop Dis. 2015;21:20. [Open Access]

Thanks to professor Lourenco for sending me his article!

A new family, genus and species of scorpion from Myanmar amber

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Andrea Rossi has recently investigated a new scorpion found in amber (burmite) from Myanmar. A new fossil family, genus and species are described.

Sucinlourencous adrianae Rossi, 2015 (Sucinlourencoidae)

Abstract:
A remarkable new family, genus and species of scorpion are described from the cretaceous burmese amber (burmite) from Myanmar. The new family Sucinlourencoidae fam. n. shows particular features that are unique among the extinct burmese families and the existing families of scorpions.

Reference:
Rossi A. A new family, genus and species of scorpion from the burmite of Myanmar (Scorpiones: Sucinlourencoidae). Arachnida - Rivista Arachnologica Italiana. 2915;1(1):3-21.

Thanks to Dr. Rossi for sending me his paper!

A new species of Centruroides from Mexico

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Rolando Teruel and co-workers have recently published a new species of Centruroides Marx, 1890 (Buthidae) from southeastern Mexico.

Centruroides poncei Teruel, Kovarik, Baldazo-Monsivais & Hoferek, 2015

The article also present data on habitat and life history for the new species.

Abstract:
The present paper deals with two species of Centruroides Marx, 1890 of the "nigrovariatus" group. One of them is described as new from two nearby localities of southwestern Oaxaca and southeastern Guerrero (in the Costa Chica region of the Pacific Coastal Plain), while the second represents a new record for Centruroides rodolfoi Santibáñez-López & Contreras Félix, 2013. After this addition, the genus is represented in the country by 41 species, eight of them belonging to the "nigrovariatus" group.

Reference:
Teruel R, Kovarik F, Baldazo Monsivaiz JG, Hoferek D. A new species of Centruroides of the "nigrovariatus" group (Scorpiones: Buthidae) from southern Mexico. Revista Iberica de Arachnologia. 2015 (26):3-14. [Subscription required for full text]

Thanks to Dr. Rolando Teruel for sending me this article!

Family Buthidae

A Revision of the Anatolian-Caucasian “Euscorpius mingrelicus Complex” and three new species

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Gioele Tropea, Ersen Yagmur and Victor Fet have recently published a revision of the Anatolian-Caucasian “Euscorpius mingrelicus Complex”, which has not been subjected to modern invesigations. As with many of the other "old"Euscorpius species, E. mingrelicus (Kessler, 1874) (Euscorpiidae) also turns out to hide several species.

These are the main conclusions of this study:

The following subspecies (to E. mingrelicus) have been given species status:

Euscorpius ciliciensis Birula, 1898 (Turkey: Konya and Mersin Provinces (eastern part of the Central Taurus Mts.))

Euscorpius phrygius Bonacina, 1980 (Turkey (northwest: Ankara, Bolu, Düzce, Karabük, and Sakarya Provinces))

Euscorpius uludagensis Lacroix, 1995 (Turkey (northwest: Bursa Province))

Euscorpius mingrelicus is now restricted to Georgia (west), Russia (southwest; Krasnodar Province near its border with northwestern Georgia) and Turkey (north).

Two subspecies are synonymized (see abstract below).

The article has habitat descriptions and pictures, and an identification key for the taxa described in the article.

Interestingly, E. ciliciensis was collected up to 2700 m, which is a record altitude for the genus.

Abstract:
The taxonomic status of the Anatolian-Caucasian subspecies of Euscorpius mingrelicus (Kessler, 1874) is reconsidered. We restrict E. mingrelicus s.str. to Georgia and Turkey, and elevate to species status three of its subspecies: E. phrygius Bonacina, 1980, stat. n., E. uludagensis Lacroix, 1995, stat. n., and E. ciliciensis Birula, 1898, stat. n. We synonymize E. mingrelicus legrandi Lacroix, 1995 with E. phrygius stat. n., and E. m. ollivieri Lacroix, 1995, with E. mingrelicus.

Reference:
Tropea G, Yagmur EA, Fet V. A Revision of the Anatolian-Caucasian “Euscorpius mingrelicus Complex” (Scorpiones: Euscorpiidae). Euscorpius. 2015 (203):1-32. [Open Access]

Thanks to Gioele Tropea for sending me the article!

Family Euscorpiidae

Two new buthids from Chad

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Wilson Lourenco has recently published two new buthids (Buthidae) from Chad.

Butheoloides vanderberghi Lourenco, 2015

Buthus labuschagnei Lourenco, 2015

Abstract:
Two new species belonging to the genera Butheoloides Hirst, 1925 (subgenus Butheoloides Hirst, 1925) and Buthus
Leach, 1815 (Scorpiones, Buthidae) are described from the Zakouma National Park, located in south-eastern Chad. The local
biotope found in the park corresponds to a more mesic environment than those found in the nearby regions, composed of dry
savannahs and the Sahel. The description of the new Butheoloides species brings further evidence to the peri-Saharan pattern of distribution presented by this genus, while the new species of Buthus is probably associated with the ‘Buthus occitanus’ from French occidental Africa (AOF), as previously reported by Vachon from this large region. This is the second record of a Buthus species from Chad.

Reference:
Lourenco WR. Deux nouvelles espèces de scorpions de la famille des Buthidae C. L. Koch, 1837 collectées dans le Parc National de Zakouma au Tchad. Revista Iberica de Arachnologia. 2015 (26):19-24. [Subscription required for full text]

Family Buthidae

A new species of Grosphus from Madagascar

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Wilson Lourenco and Lucienne Wilme have investigated scorpion materials collected in the Makay Mountain range in Madagascar. One new species of Grosphus Simon, 1880 (Buthidae) has been described.

Grosphus makay Lourenco & Wilme, 2015

The taxonomic history of Grosphus limbatus (Pocock, 1889) is also discussed.

Abstract:
New considerations regarding Grosphus limbatus (Pocock, 1889), described from an undefined locality in Madagascar, are proposed. One new species, Grosphus makay sp. n., closely allied to G. limbatus, is described from the Makay sandstone mountain range, Atsimo-Andrefana Region (ex Province of Toliara) in south-western Madagascar. The description of the new species brings further evidence of the existence of micro-endemic and vicariant populations within the Malagasy scorpion fauna.

Reference:
Lourenco WR, Wilme L. Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species. Revista Iberica de Arachnologia. 2015 (26):55-61. [Subscription required for full text]

Family Buthidae

A new species of Hottentotta from India

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Wilson Lourenco has recently published a paper on the buthid genus Hottentotta Birula, 1908, describing a new species from southern India.

Hottentotta reddyi Lourenco, 2015

The taxonomic position of Hottentotta and its relation to Mesobuthus Vachon, 1950 is discussed.

Abstract:
A new species of scorpion, Hottentotta reddyi sp. n. (Buthidae), is described. The type materials was collected Thorn Scrub vegetation formation, NW of Salem, in the state of Tamil Nadu, in the south of India. This new species may be the first official precise record of a Hottentotta species for this region of India. Some comments are also added about the characters used in the diagnosis of the genus Hottentotta, and their possible need for the definition of subgenera or, at least, groups of species.

Reference:
Lourenco WR. New considerations on the genus Hottentotta Birula, 1908 (Scorpiones: Buthidae) with the description of one new species from the south of India. Arachnida - Rivista Arachnologica Italiana. 2015;1(2):37-49.

Thanks to professor Lourenco for sending me his article!

Family Buthidae

An updated checklist of the scorpion fauna of Ecuador and a review on their medical importance

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Gabriel Brito and Adolfo Borges have recently published an interesting article on the scorpion fauna of Ecuador. Ecuador is a hotspot for scorpion biodiversity and the authors present an updated checklist of the scorpion fauna of this country. Details of distribution and habitat are also presented.

The article also discuss the medical importance of the scorpion fauna of Ecuador.

Abstract:
Ecuador harbors one of the most diverse Neotropical scorpion faunas, hereby updated to 47 species contained within eight genera and five families, which inhabits the “Costa” (n = 17), “Sierra” (n = 34), “Oriente” (n = 16) and “Insular” (n = 2) biogeographical regions, corresponding to the western coastal, Andean, Amazonian, and the Galápagos archipelago regions, respectively. The genus Tityus Koch, in the family Buthidae, responsible for severe/fatal accidents elsewhere in northern South America and the Amazonia, is represented in Ecuador by 16 species, including T. asthenes, which has caused fatalities in Colombia and Panama, and now in the Ecuadorian provinces of Morona Santiago and Sucumbíos. Underestimation of the medical significance of scorpion envenoming in Ecuador arises from the fact that Centruroides margaritatus (Gervais) (family Buthidae) and Teuthraustes atramentarius Simon (family Chactidae), whose venoms show low toxicity towards vertebrates, frequently envenom humans in the highly populated Guayas and Pichincha provinces. This work also updates the local scorpion faunal endemicity (74.5 %) and its geographical distribution, and reviews available medical/biochemical information on each species in the light of the increasing problem of scorpionism in the country. A proposal is hereby put forward to classify the Ecuadorian scorpions based on their potential medical importance.

Reference:
Brito G, Borges A. A checklist of the scorpions of Ecuador (Arachnida: Scorpiones), with notes on the distribution and medical significance of some species. J Venom Anim Toxins Incl Trop Dis. 2015;21:23. [Open Access]

Thanks to Dr. Borges for sending me this article!

A new species of Compsobuthus from Armenia

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Wilson Lourenco and Elise Ann-Leguin have recently discovered a new species of Compsobuthus Vachon, 1949 from Armenia.

Compsobuthus armenicus Lourenco & Leguin, 2015

Abstract:
Two buthid species previously recorded from Armenia are confirmed: Androctonus crassicauda (Olivier, 1807) and Mesobuthus eupeus (C. L. Koch, 1839). Hottentotta saulcyi (Simon, 1880), is recorded for the first time for this country. A new species of Compsobuthus Vachon, 1949 is described on the basis of two adult females and three juveniles. From the absence of outer denticles on the movable finger of the pedipalps, the new species is placed in the acutecarinatus group. The new species of Compsobuthus represents the first record of this genus for Armenia.

Reference:
Lourenco WR, Leguin E-A. A new species of Compsobuthus Vachon, 1949 (Scorpiones: Buthidae) from Armenia. Zoology in the Middle East. 2015:1-5. [Subscription required for full text]

Thanks to professor Lourenco for sending me this article!

Family Buthidae

A new species of Uroplectes from South Africa

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Lorenzo Prendini has recently described a new species of Uroplectes Peters, 1861 (Buthidae) from the Karoo Biome in the Northern Cape and Western Cape provinces of South Africa.

Uroplectes ansiedippenaarae Prendini, 2015

The new species is one of the smallest species in Africa with adult size between 16 and 20 mm.

Abstract:
The scorpion fauna of southern Africa is very diverse, especially in the arid western half of the subcontinent. New species continue to be discovered as the region is surveyed with ultraviolet light detection methods. The present contribution describes Uroplectes ansiedippenaarae sp. n., which is endemic to the Succulent Karoo Biome in the Northern Cape and Western Cape provinces of South Africa. The new species appears to be most closely related to U. variegatus (C.L. Koch, 1844), which is endemic to the Fynbos Biome in the Western Cape Province. Uroplectes ansiedippenaarae sp. n. is the smallest species of Uroplectes Peters, 1861, and among the smallest scorpion species in southern Africa, with adults ranging from 16–20 mm in total length. The addition of this new species raises the number of Uroplectes species and subspecies in South Africa to 19, and the number of endemics to 10.

Reference:
Prendini L. A remarkably small species of Uroplectes Peters, 1861 (Scorpiones: Buthidae), endemic to the Succulent Karoo of South Africa. African Invertebrates. 2015;56(2):499-513. [Open Access]

Thanks to Rolando Teruel for informing me about this article!

Family Buthidae

New fossil scorpion from Mexican amber

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Francisco Riquelme and co-workers have recently described a new fossil scorpion from amber found in the Chipas Highlands in southern Mexico.

Tityus apozonelli Riquelme, Villegas & González, 2015 (Buthidae)

Please note that the species is extinct, even though it is placed in an extant genus.

Abstract:
A new species of scorpion is described based on a rare entire adult male preserved in a cloudy amber from Miocene rocks in the Chiapas Highlands, south of Mexico. The amberbearing beds in Chiapas constitute a Conservation Lagerstätte with outstanding organic preservation inside plant resin. The new species is diagnosed as having putative characters that largely correspond with the genus Tityus Koch, 1836 (Scorpiones, Buthidae). Accordingly, it is now referred to as Tityus apozonalli sp. nov. Its previously unclear phylogenetic relationship among fossil taxa of the family Buthidae from both Dominican and Mexican amber is also examined herein. Preliminarily results indicate a basal condition of T. apozonalli regarding to Tityus geratus Santiago-Blay and Poinar, 1988, Tityus (Brazilotityus) hartkorni Lourenço, 2009, and Tityus azari Lourenço, 2013 from Dominican amber, as was Tityus (Brazilotityus) knodeli Lourenço, 2014 from Mexican amber. Its close relationships with extant Neotropic Tityus-like subclades such as ‘Tityus clathratus’ and the subgenus Tityus (Archaeotityus) are also discussed. This new taxon adds to the knowledge of New World scorpions from the Miocene that are rarely found trapped in amber.

Reference:
Riquelme F, Villegas-Guzman G, Gonzalez-Santillan E, Cordova-Tabares V, Francke OF, Piedra-Jimenez D, et al. New Fossil Scorpion from the Chiapas Amber Lagerstatte. PLoS One. 2015;10(8):e0133396. [Open Access]

Thanks to Oscar Francke for sending me this article!

A new species of Buthoscorpio from India

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Aswathi, Sureshan and Lourenco has recently published the discovery of a new species of Buthoscorpio Werner, 1936 (Buthidae). This is a rare genus and the new species is the fifth in the genus (three in India and one in Sri Lanka).

Buthoscorpio chinnarensis Awathi, Sureshan & Lourenco, 2015

Abstract:
A new scorpion species of the genus Buthoscorpio Werner, 1936 is described from Chinnar Wildlife Sanctuary, Idukki District, Kerala, India. The new species, Buthoscorpio chinnarensis can be distingued from the other congeners by having (1) carapace with scattered granulation on lateral portions; (2) median eyes situated anteriorly in the ratio 1:2; (3) prominent tubercles present at the basal portion of the pedipalp femur; (4) marginal lamellae of pectines composed of three pieces and middle lamellae comprises seven pieces of sclerites; (5) dorsal carinae present on metasomal segments III–V; (6) subaculear tubercle absent on telson; (7) Pectinal teeth count 14/16 (female paratypes with 17/17 and 16/17 respectively); (8) Metasomal segments I–IV wider than long, segment V longer than wide; (9) arrangement of lateral eyes. After the description of this new species, the total number of known species in the genus is raised to five; four in India and one in Sri Lanka.

Reference:
Aswathi K, Sureshan PM, Lourenco WR. A new scorpion of the genus Buthoscorpio Werner, 1936 (Scorpiones: Buthidae) from Kerala, India. Taprobanica. 2015;7(4):213-8. [Subscription required for full text]

Thanks to professor Lourenco for sending me this article!

Family Buthidae

A new species of Vachonus from India

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Wilson Lourenco has discovered a new species of Vachonus Tikader & Bastawade, 1983 (Buthidae) from India.

Vachonus inexpectatus Lourenco, 2015

Abstract:
Comments are proposed on the enigmatic genus Vachonus Tikader et Bastawade, 1983. Some new characters bring further evidence to support the validity of this genus. One new species is described on the basis of two adult female specimens collected in the State of Gujarat, E of Ghandinagar, in India.

Reference:
Lourenco WR. New considerations on the enigmatic genus Vachonus Tikader et Bastawade, 1983, with the description of a new species from India (Scorpiones: Buthidae). Arachnida - Rivista Arachnologica Italiana. 2015;1(3):2-9.

Thanks to professor Lourenco for sending me this article!

Family Buthidae

A new Euscorpius species from Turkey

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Ersen Yagmur and Gioele Tropea have recently described a new species of Euscorpius Thorell, 1876 (Euscorpiidae) from southwestern Turkey.

Euscorpius arikani Yagmur & Tropea, 2015

The paper has an identification key for the Euscorpius species in the former "Euscorpius mingrelicus species complex" in Georgia and Turkey.

Abstract:
A new scorpion species, Euscorpius arikani sp. n., is described based on specimens collected from Antalya Province, in southwestern Turkey. It is characterized by the trichobothrial series Pe-em = 3, small size, and light to medium brown-reddish colour.

Reference:
Yagmur EA, Tropea G. A new species of Euscorpius Thorell, 1876, from southwestern Turkey (Scorpiones: Euscorpiidae). Arachnida - Rivista Arachnologica Italiana. 2015;1(3):14-26.

Thanks to Ersen Yagmur for sending me this article!

Family Euscorpiidae

Two new fossil species from Cretaceous Burmese amber

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Wilson Lourenco has recently described a couple new fossil species of Betaburmesebuthus Lourenco, 2015 (Arachaeobuthidae) from Cretaceous Burmese amber.

Betaburmesebuthus muelleri Lourenco, 2015

Betaburmesebuthus bidentatus Lourenco, 2015 

Abstract:
The study of three new scorpion specimens from Cretacous amber of Myanmar (Burma) leads to the description of two new species in the genus Betaburmesebuthus Lourenco, 2015 bringing further support for the validity of this genus. To date, five species have been described in the subfamily Palaeoburmesebuthinae Lourenco, 2015: two in the genus Palaeoburmesebuthus Lourenco, 2002 and three in the genus Betaburmesebuthus.This further study attests to a considerable diversity in the Burmese amber -producing forests.

Reference:
Lourenco WR. New contribution to the knowledge of Cretaceous Burmese amber scorpions: descrptions of two new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Arachnida - Rivista Arachnologica Italiana. 2015;1(3):27-36.

Thanks to professor Lourenco for sending me his article!
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